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Fish erythrocyte sizes

NOTE: These are size measurements for fish erythrocytes (red blood cells).  Erythrocytes in fishes are elliptical, and as such two different diameters are provided: CLD ("cell long diameter") and CSD ("cell short diameter").  NLD ("nucleus long diameter") and NSD ("nucleus short diameter") are also provided where available.  In both cases, the diameters were measured from dry smears.  These were used to calculate dry cell area (CA) and dry nucleus area (NA) using the equation for elliptical area: CA = pi * (CLD/2) * (CSD/2).  In some cases, area is measured directly using image analysis with a calibration slide, so diameters are not available.  MCV ("mean corpuscular volume") is wet volume and is usually measured with a Coulter counter.  These different measurements (e.g., CA and MCV) should not be compared to each other, so pick only one for any analyses you wish to do, or run them separately.  Diameters are in µm, areas are in µm2, and volumes are in µm3. 

Species and taxonomic information   
CLD   
CSD  
CA      
Ref  
NLD  
NSD  
NA     
Ref  
MCV  
Ref   
SUPERCLASS AGNATHA
CLASS CEPHALASPIDOMORPHI
ORDER Petromyzontiformes
Family Petromyzontidae
Geotria australis 12.8 12.8 128.68 11



370 11
Lampetra fluviatilis 11.6 11.6 105.68 11



189 11
" " 11.9 11.9 111.27 1
Lampetra planeri 11.9 11.9 111.27 1
Petromyzon marinus 11.9 11.9 111.27 1 4.0 4.0 12.37 1 534 13
" " 14.3 14.3 160.61 6



710 6
CLASS MYXINI
ORDER Myxiniformes
Family Myxinidae
Myxine glutinosa 26.2 18.0 370.39 3



1530 3
Myxus petardi 10.2 7.0 56.08 2 3.5 2.5 6.87 2
SUPERCLASS GNATHOSTOMATA
CLASS CHONDRICHTHYES
SUBCLASS ELASMOBRANCHII
ORDER Carcharhiniformes
Family Carcharhinidae
Carcharhinus brachyurus

175.36 14

31.92 14
Carcharhinus leucas 17.5 12.7 174.55 10
Carcharhinus maculipinnis 15.1 10.4 123.34 13 5.8 4.3 19.59 13 739 13
Carcharhinus melanopterus

143.91 14

27.33 14
Carcharhinus obscurus 19.0 13.5 201.45 4 7.3 6.5 37.27 4
" " 16.3 14.9 190.75 10
Carcharhinus plumbeus 16.7 15.5 203.30 10
Galeocerdo cuvier 21.9 17.9 307.88 10
Negaprion brevirostris 15.3 12.8 153.81 10
Prionace glauca 21.6 15.4 261.25 13 7.9 6.8 42.19 13 800 13
Rhizoprionodon terraenovae 19.3 16.2 245.56 10
Family Scyliorhinidae
Scyliorhinus canicula 25.4 16.5 328.60 1 8.5 6.4 42.23 1
Scyliorhinus stellaris 25.4 14.3 285.15 1
Family Sphyrnidae
Sphyrna lewini

229.69 14

40.63 14
Sphyrna mokarran 17.0 12.3 164.23 10
Sphyrna tiburo 17.5 12.7 174.55 10
Sphyrna tudes 18.0 12.5 176.71 2 7.0 5.5 30.24 2
Sphyrna zygaena 14.0 10.6 116.55 6



526 6
Family Triakidae
Mustelus canis 16.2 10.8 137.41 4 6.5 5.2 26.55 4 546 5
ORDER Lamniformes
Family Lamnidae
Isurus oxyrinchus

231.39 14

78.07 14 727 13
Lamna nasus 27.5 16.9 365.99 1
ORDER Orectolobiformes
Family Ginglymostomatidae
Ginglymostoma cirratum 18.4 16.0 231.22 10
Nebrius ferrugineus

250.48 14

64.46 14
Family Hemiscylliidae
Chiloscyllium punctatum

261.88 14

50.01 14
Chiloscyllium sp. 23.0 13.5 243.87 2 9.0 7.0 49.48 2
Hemiscyllium ocellatum

195.86 14

40.31 14
Family Orectolobidae
Orectolobus ornatus

318.06 14

51.17 14
ORDER Rajiformes
Family Dasyatidae
Dasyatis bennetti

176.42 14

33.16 14
Dasyatis centroura 19.7 13.8 213.52 7 8.1 6.9 43.90 7
" " 21.1 14.8 245.26 5
Dasyatis kuhlii 18.0 14.0 197.92 2 7.0 5.5 30.24 2
Himantura signifer

166.70 14

39.57 14
Family Rajidae
Bathyraja parmifera

225.29 14

59.84 14
Dipturus batis 26.2 17.5 359.02 1
Dipturus chilensis 21.3 14.5 242.57 4 8.1 6.0 38.17 4 727 5
" " 28.4 17.5 390.34 3



1274 3
Dipturus laevis 23.3 16.9 309.91 1
Leucoraja erinacea 20.3 12.9 205.67 7 8.4 6.1 40.24 7
" " 24.3 13.9 265.28 3



778 3
Leucoraja ocellata

246.45 14

39.01 14
Raja clavata 23.3 15.9 290.54 1
Raja eglanteria 23.3 14.1 258.03 3



766 3
" " 23.6 16.4 303.98 13 6.4 5.1 25.64 13 1095 13
" " 24.3 14.3 272.92 4 6.7 5.4 28.42 4 864 5
" " 25.5 14.6 292.40 10
Raja montagui 22.0 14.1 243.63 6



641 6
Family Rhinobatidae
Aptychotrema rostrata

236.70 14

39.20 14
ORDER Squaliformes
Family Dalatiidae
Centroscymnus coelolepis

599.60 14

127.09 14
Centroscymnus crepidater

609.98 14

147.57 14
Centroscymnus owstoni

591.67 14

124.07 14
Centroscymnus plunketi

482.56 14

157.33 14
Etmopterus brachyurus

556.70 14

144.96 14
Etmopterus granulosus

492.36 14

134.46 14
Oxynotus bruniensis

639.02 14

110.68 14
Oxynotus centrina 31.8 25.4 633.38 1
Family Squalidae
Squalus acanthias 23.1 15.4 279.40 13 8.7 5.7 38.95 13 750 13
" " 22.1 15.9 275.86 1 8.5 6.4 42.23 1
" " 23.7 17.0 316.44 3



947 3
" "

326.42 14

55.17 14
ORDER Squatiniformes
Family Squatinidae
Squatina australis

497.46 14

142.49 14
Squatina squatina 24.6 17.5 337.45 1
ORDER Torpediniformes
Family Torpedinidae
Torpedo nobiliana 29.8 23.2 542.99 6



1593 6
Torpedo torpedo 31.8 25.4 633.38 1
CLASS OSTEICHTHYES
SUBCLASS ACTINOPTERYGII
INFRACLASS Chondrostei
ORDER Acipenseriformes
Family Acipenseridae
Acipenser brevirostrum

192.74 14

43.82 14
" " (triploid)

234.10 14

64.60 14
Acipenser oxyrhynchus

96.47 14

18.88 14
" " (triploid)

141.23 14

23.63 14
Acipenser sturio 14.0 9.7 106.66 5



506 5
" " 15.9 9.5 118.79 1
Family Polypteridae
Polypterus palmas

158.87 14

34.72 14
INFRACLASS Neopterygii
SUPERORDER Teleostei
ORDER Anguilliformes
Family Anguillidae
Anguilla anguilla 14.6 8.9 102.17 1
Anguilla bicolor bicolor 14.3 8.5 95.05 1
Anguilla marmorata 13.4 8.8 92.20 1
Anguilla rostrata 13.0 8.0 81.68 3



156 3
" " 11.5 7.6 68.64 7 3.8 3.2 9.55 7
" "

101.88 14

16.71 14
Anguilla sp. 12.3 8.1 78.25 5
Family Chlopsidae
Ophichthus cephalozona

99.34 14

20.24 14
Family Congridae
Conger conger 11.1 8.5 73.89 1
Family Muraenesocidae
Muraenesox cinereus

87.77 14

22.62 14
Family Muraenidae
Sideria picta

123.73 14

18.99 14
Family Ophichthidae
Ophichthus cruentifer 10.9 7.7 65.92 7 4.0 3.3 10.37 7
Family Synaphobranchidae
Diastobranchus capensis

201.00 14

26.75 14
ORDER Atheriniformes
Family Atherinidae
Odontesthes argentinensis 12.0 7.5 70.69 9 5.0 3.0 11.78 9
ORDER Aulopiformes
Family Synodontidae
Synodus sageneus

67.32 14

16.32 14
ORDER Batrachoidiformes
Family Batrachoididae
Opsanus tau 17.1 11.5 154.45 13 5.9 4.3 19.93 13 800 13
" " 14.9 12.9 150.96 7 5.8 5.2 23.69 7
ORDER Beloniformes
Family Belonidae
Belone belone 11.1 8.5 73.89 1
Platybelone argalus platyura

74.71 14

18.20 14
ORDER Beryciformes
Family Diretmidae
Diretmichthys parini

106.75 14

19.01 14
ORDER Characiformes
Family Characidae
Colossoma macropomum

71.90 14

18.08 14
Hemigrammus caudovittatus 16.5 9.0 116.63 9 6.0 4.5 21.21 9
ORDER Clupeiformes
Family Chirocentridae
Chirocentrus dorab

57.50 14

12.49 14
Family Clupeidae
Alosa fallax 11.1 8.5 73.89 1
Brevoortia tyrannus 9.8 7.3 56.19 4 4.6 3.2 11.56 4
Clupea harengus 9.5 7.1 53.46 1
Nematalosa come

62.94 14

10.85 14
Nematalosa erebi 12.2 8.0 76.65 2 4.8 2.3 8.67 2
Sardina pilchardus 11.1 7.9 69.27 1
Sardinella gibbosa

59.94 14

15.09 14
Sprattus sprattus 9.5 7.1 53.46 1
Family Engraulidae
Engraulis anchoita 12.0 7.5 70.69 9 5.0 4.0 15.71 9
Engraulis encrasicolus 11.1 8.5 73.89 1
ORDER Cypriniformes
Family Balitoridae
Barbatula barbatula 11.1 7.9 69.27 1
Family Catostomidae
Catostomus catostomus catostomus

137.82 14

26.57 14
Catostomus commersonii

105.44 14

21.97 14
Family Cyprinidae
Abramis brama 10.6 7.1 59.39 1
Carassius auratus auratus 12.5 8.5 83.45 9 5.5 4.0 17.28 9
" " 14.3 9.0 100.97 1
" "

82.69 14

12.72 14
Cyprinus carpio 11.9 7.4 68.99 1 4.0 3.2 9.90 1
Epalzeorhynchos bicolor

53.20 14

12.65 14
Epalzeorhynchos frenatum

55.17 14

14.73 14
Gobio gobio 11.9 8.8 81.92 1
Labeo chrysophekadion

60.03 14

11.98 14
Leuciscus cephalus 11.9 7.1 66.82 1
Phoxinus eos-neogaeus

132.19 14

23.12 14
Phoxinus neogaeus

94.19 14

15.91 14
Phoxinus phoxinus 12.7 8.8 87.36 1
Rutilus rutilus 11.9 8.5 79.19 1
Scardinius erythrophthalmus 12.7 7.9 79.17 1
Semotilus corporalis

74.07 14

14.99 14
Tinca tinca 13.4 10.6 111.41 1 3.2 2.6 6.60 1
Family Gyrinocheilidae
Gyrinocheilus aymonieri

40.43 14

7.57 14
ORDER Cyprinodontiformes
Family Anablepidae
Jenynsia lineata 10.5 6.5 53.60 9 4.5 3.0 10.60 9
Family Aplocheilidae
Pachypanchax playfairii

37.63 14

8.58 14
Family Poeciliidae
Poecilia mexicana

60.97 14

13.40 14
ORDER Esociformes
Family Esocidae
Esox lucius 12.7 7.1 71.27 1 4.8 3.2 11.88 1
" "

73.84 14

16.63 14
Esox niger

75.30 14

13.72 14
ORDER Gadiformes
Family Gadidae
Boreogadus saida

89.24 14

12.65 14
Gadus morhua

76.51 14

17.33 14
Gadus morhua 12.2 9.0 86.24 3



186 3
" " 11.9 7.1 66.82 1
Melanogrammus aeglefinus 10.3 7.9 64.37 1
Merlangius merlangus 10.3 7.1 57.94 1
Pollachius virens 11.4 8.1 72.52 3



143 3
" " 11.9 8.5 79.19 1
Trisopterus luscus 10.3 7.9 64.37 1
Family Lotidae
Ciliata mustela 10.6 7.9 65.98 1
Gaidropsarus ensis

86.95 14

13.81 14
Family Macrouridae
Caelorinchus innotabilis

67.35 14

12.35 14
Caelorinchus maurofasciatus

67.07 14

12.82 14
Coryphaenoides serrulatus

105.73 14

20.05 14
Macrourus berglax

62.17 14

12.82 14
Mesobius antipodum

112.19 14

20.07 14
Family Merlucciidae
Merluccius bilinearis 11.1 5.9 51.44 3



254 3
Merluccius hubbsi 15.0 6.5 76.58 9 6.0 3.0 14.14 9
Merluccius merluccius 11.9 8.5 79.19 1
Family Moridae
Halargyreus johnsonii

102.21 14

18.26 14
Family Phycidae
Urophycis tenuis

72.90 14

17.76 14
" " 13.7 8.0 86.08 3



242 3
ORDER Gasterosteiformes
Family Gasterosteiidae
Gasterosteus aculeatus aculeatus 9.5 8.5 63.35 1
Pungitius pungitius pungitius 9.5 8.5 63.35 1
ORDER Gymnotiformes
Family Electrophoridae
Electrophorus electricus 14.6 9.8 111.73 1
Family Rhamphichthyidae
Rhamphichthys rostratus

41.83 14

14.79 14
ORDER Lophiiformes
Family Lophiidae
Lophius americanus

84.21 14

17.70 14
Lophius piscatorius 11.6 9.2 83.82 13 5.6 4.2 18.47 13 215 13
" " 13.8 9.3 100.80 3



167 3
" " 12.6 10.8 106.88 7 5.2 4.7 19.20 7
" " 13.0 10.3 105.16 6



241 6
" " 13.4 9.5 100.30 1
Family Tetrabranchiidae
Tetrabrachium ocellatum

84.62 14

18.19 14
ORDER Myctophiformes
Family Neoscopelidae
Neoscopelus macrolepidotus

153.71 14

29.93 14
ORDER Osmeriformes
Family Galaxiidae
Galaxias olidus 11.0 9.0 77.75 2 5.5 3.5 15.12 2
Family Osmeridae
Osmerus eperlanus 11.1 8.5 73.89 1 4.0 2.9 8.91 1
Osmerus mordax

61.20 14

17.10 14
ORDER Perciformes
Family Acanthuridae
Acanthurus grammoptilus

49.53 14

11.76 14
Zebrasoma scopas

58.40 14

12.53 14
Family Apogonidae
Apogon cookii

66.93 14

13.62 14
Apogon endetakaena

70.61 14

12.99 14
Apogon guamensis

70.03 14

13.96 14
Apogon kallopterus

66.46 14

14.70 14
Apogon savayensis

71.44 14

17.99 14
Family Ammodytidae
Ammodytes tobianus 12.7 7.1 71.27 1
Family Belontidae
Betta splendens

56.86 14

8.92 14
Family Blenniidae
Crossosalarias macrospilus

65.14 14

10.73 14
Ecsenius mandibularis

50.66 14

13.30 14
Ecsenius yaeyamaensis

48.38 14

11.25 14
Lipophrys pholis 10.3 8.5 68.66 1
Petroscirtes fallax

55.97 14

9.25 14
Petroscirtes lupus

60.30 14

10.47 14
Petroscirtes mitratus

54.80 14

7.06 14
Plagiotremus rhinorhynchos

59.77 14

13.80 14
Family Caesionidae
Caesio cuning

67.28 14

7.56 14
Family Callionymidae
Callionymus limiceps

53.63 14

14.95 14
Callionymus lyra 11.1 8.5 73.89 1
Family Carangidae
Carangoides fulvoguttatus

48.63 14

9.96 14
Carangoides gymnostethus

45.45 14

8.89 14
Seriola hippos

46.03 14

10.25 14
Seriola lalandi 10.5 7.0 57.73 2 4.5 2.5 8.84 2
Trachurus trachurus 12.7 7.1 71.27 1
Ulua aurochs

38.74 14

9.56 14
Family Chaetodontidae
Chaetodon lunulatus

54.87 14

12.23 14
Chaetodon rainfordi

44.22 14

11.64 14
Family Channidae
Channa gachua







133 12
Channa punctata







130 12
Channa striata







126 12
Family Coryphaenidae
Coryphaena hippurus

42.56 14

11.06 14
Family Cryptacanthodidae
Cryptacanthodes maculatus 13.5 9.0 95.43 3



200 3
Family Drepaneidae
Drepane punctata

54.62 14

10.43 14
Family Echeneidae
Echeneis naucrates 11.8 7.0 64.87 5



111 5
" " 9.0 7.0 49.48 2 4.5 2.5 8.84 2
" "

42.16 14

6.79 14
Family Gerreidae
Gerres subfasciatus

46.74 14

7.54 14
Family Gobiidae
Cryptocentrus leptocepgus

79.81 14

19.58 14
Gobiodon citrinus

58.22 14

16.48 14
Istigobius rigilius

100.36 14

30.76 14
Neogobius melanostomus

80.93 14

20.87 14
Oplopomus oplopomus

76.04 14

20.00 14
Paragobiodon xanthosoma

54.76 14

13.89 14
Signigobius biocellatus

68.03 14

13.17 14
Valenciennea longipinnis

66.67 14

17.80 14
Family Haemulidae
Diagramma labiosum

49.30 14

9.67 14
Plectorhinchus sp. 9.0 7.0 49.48 2 4.5 2.5 8.84 2
Family Labridae
Choerodon cephalotes

59.39 14

14.44 14
Choerodon fasciatus

73.19 14

12.36 14
Coris batuensis

46.54 14

12.49 14
Halichoeres biocellatus

67.03 14

13.55 14
Novaculichthys taeniourus

46.33 14

8.48 14
Tautoga onitis 10.5 7.3 60.20 7 4.1 3.0 9.66 7
" " 10.8 7.8 66.16 13 4.4 3.5 12.10 13 102 13
" " 10.8 7.3 61.92 4 4.3 3.2 10.81 4
Family Leiognathidae
Leiognathus decorus

40.37 14

11.43 14
Family Lethrinidae
Lethrinus atkinsoni

63.95 14

12.54 14
Lethrinus miniatus 9.0 8.0 56.55 2 5.5 3.5 15.12 2
Lethrinus nebulosus

65.57 14

13.00 14
Lethrinus rubrioperculatus

71.09 14

13.65 14
Family Lutjanidae
Aprion virescens

62.43 14

11.69 14
" " (tetraploid?)

78.22 14

17.17 14
Lutjanus carponotatus

62.37 14

11.19 14
Lutjanus fluviflamma

56.36 14

11.26 14
Lutjanus sebae

57.79 14

11.62 14
Lutjanus vitta

57.70 14

9.47 14
Family Moronidae
Dicentrarchus labrax 9.5 7.9 59.39 1
Morone americana 10.3 7.2 58.25 3



113 3
Morone saxatilis

57.33 14

12.23 14
Family Mugilidae
Liza ramada 10.6 7.9 65.98 1
Liza vaigiensis

52.82 14

7.78 14
Myxus elongatus

51.14 14

8.91 14
Family Mullidae
Mulloidichthys vanicolensis

42.44 14

11.15 14
Mullus barbatus 11.1 7.9 69.27 1
Mullus surmuletus 11.1 8.5 73.89 1
Upeneichthys lineatus

51.11 14

11.86 14
Family Nemipteridae
Scolopsis monogramma

51.41 14

13.47 14
Scolopsis vosmeri 10.5 7.0 57.73 2 6.5 3.0 15.32 2
Family Nototheniidae
Dissostichus mawsoni

107.51 14

15.67 14
Trematomus bernacchii

89.19 14

13.50 14
Family Opistognathidae
Opistognathus sp.

95.06 14

18.08 14
Family Percidae
Gymnocephalus cernuus 10.3 8.5 68.63 1
Perca fluviatilis 12.1 9.0 85.48 1 3.4 2.9 7.67 1
Family Pinguipedidae
Parapercis cylindrica

42.96 14

9.18 14
Parapercis hexophtalma

58.70 14

10.37 14
Family Pomacanthidae
Centropyge bicolor

48.13 14

11.60 14
Family Pomacentridae
Amphiprion akindynos

57.56 14

14.55 14
Amphiprion clarkii

44.92 14

12.89 14
Chromis analis

50.29 14

8.50 14
Chromis viridis

59.50 14

9.24 14
Chrysiptera cyanea (male)

66.12 14

10.11 14
" " (female)

54.00 14

9.11 14
Dascyllus aruanus

55.70 14

9.00 14
Dischistodus prosopotaenia

55.58 14

14.21 14
Hemiglyphidodon plagiometopon

72.42 14

16.94 14
Pomacentrus nagasakiensis

59.95 14

11.19 14
Premnas biaculeatus

59.77 14

16.38 14
Family Priacanthidae
Priacanthus tayenus

46.47 14

10.98 14
Family Rachycentridae
Rachycentron canadus 8.0 6.0 37.70 13 3.7 2.7 7.85 13 259 13
Family Scaridae
Scarus schlegeli

67.39 14

19.59 14
Family Scatophagidae
Scatophagus multifasciatus

55.01 14

10.83 14
Family Scombridae
Sarda australis

56.08 14

13.84 14
Scomber scombrus 12.5 8.3 81.49 3



146 3
" " 12.0 8.2 77.28 5



118 5
" " 11.1 7.9 69.27 1
Thunnus alalunga

75.31 14

12.99 14
Family Serranidae
Aulacocephalus temminckii

49.70 14

16.04 14
Cephalopholis cyanostigma

82.95 14

17.53 14
Epinephelus cyanopodus

64.82 14

13.91 14
Epinephelus merra

73.76 14

14.42 14
Epinephelus spilotoceps

56.34 14

11.18 14
Plectropomus leopardus

47.12 14

12.66 14
Family Siganidae
Siganus doliatus

51.96 14

11.34 14
Siganus fuscescens

45.05 14

12.20 14
Family Sillaginidae
Sillaginodes punctata

50.68 14

10.90 14
Sillago analis

32.34 14

9.01 14
Family Sparidae
Pagellus bogaraveo 11.1 7.9 69.27 1
Stenotomus chrysops 8.8 7.5 51.84 13 4.0 3.0 9.42 13 210 13
Family Sphyraenidae
Sphyraena obtusata

41.97 14

10.01 14
Family Terapontidae
Leiopotherapon unicolor 6.5 6.5 33.18 2 3.5 3.5 9.62 2
Pelates quadrilineatus

52.53 14

8.83 14
Terapon jarbua

46.60 14

9.96 14
Terapon puta

51.28 14

8.33 14
Family Trachinidae
Trachinus draco 10.6 8.5 70.38 1
Family Tripterygiidae
Ucla xenogrammus

60.13 14

11.75 14
Family Uranoscopidae
Kathetostoma canaster

104.61 14

12.71 14
Family Xiphiidae
Xiphias gladius 11.7 8.3 76.27 4 4.0 3.4 10.68 4
" " 10.3 6.3 51.37 1
Family Zoarcidae
Bothrocara pusillum

56.34 14

15.50 14
Gymnelus viridis

97.92 14

14.90 14
Lycodichthys dearborni

119.11 14

18.51 14
Zoarces americanus

50.65 14

12.61 14
ORDER Pleuronectiformes
Family Paralichthyidae
Pseudorhombus jenynsii

45.61 14

7.63 14
Paralichthys lethostigma 8.8 6.3 43.54 13 3.8 2.6 7.76 13 120 13
Family Pleuronectidae
Glyptocephalus cynoglossus 12.5 7.1 69.70 3



141 3
Hippoglossus hippoglossus

70.52 14

16.93 14
Lepidopsetta bilineata

44.38 14

9.97 14
Limanda aspera

67.10 14

12.42 14
Limanda ferruginea 10.3 7.7 62.29 3



118 3
Limanda ferruginea

59.03 14

13.72 14
Limanda limanda 8.8 7.9 54.60 1
Platichthys flesus 9.5 8.5 63.35 1
Pleuronectes americanus

50.45 14

8.28 14
Pleuronectes platessus 9.5 8.5 63.35 1
Pseudopleuronectes americanus







142 8
Reinhardtius evermanni

54.06 14

12.27 14
Family Psettodidae
Psettodes erumei

68.22 14

10.03 14
Family Scophthalmidae
Scophthalmus maximus

53.80 14

13.44 14
Scophthalmus rhombus 10.3 7.9 64.37 1
Family Soleidae
Solea solea 10.6 7.9 65.98 1
Soleichthys heterorhinos

40.71 14

10.16 14
ORDER Salmoniformes
Family Salmonidae
Coregonus clupeaformis

118.22 14

23.95 14
Oncorhynchus keta

131.08 14

35.16 14
Oncorhynchus mykiss







274 8
" "

101.65 14

19.19 14
Oncorhynchus tschawytscha

109.19 14

26.35 14
Prosopium cylindraceum

105.51 14

19.47 14
Salmo salar 16.7 10.3 135.16 1
" " (diploid)

135.25 14

25.33 14
" " (triploid)

196.97 14

42.14 14
Salmo trutta 16.7 8.8 114.65 1
Salvelinus alpinus (from Labrador)

164.56 14

30.27 14
" " (from Somerset Is.)

135.35 14

28.09 14
Salvelinus fontinalis 17.5 11.1 152.34 1
" "

137.53 14

35.09 14
Salvelinus namaycush 16.0 11.3 142.00 13 6.5 4.4 22.46 13
" " 16.7 8.8 114.65 1
" "

150.45 14

24.55 14
Thymallus arcticus 12.2 9.1 87.19 13 5.4 3.8 16.12 13
" "

116.93 14

22.40 14
Thymallus thymallus 15.1 8.8 103.76 1
ORDER Scorpaeniformes
Family Agonidae
Podothecus acipenserinus

53.99 14

16.33 14
Family Cottidae
Cottus gobio 12.7 8.8 87.36 1
Icelus spatula

67.11 14

9.93 14
Myoxocephalus octodecemspinosus 11.6 8.9 81.08 3



175 3
Myoxocephalus scorpius 12.4 9.5 92.52 3



214 3
" " 12.7 8.8 87.36 1
Taurulus bubalis 12.7 8.8 87.36 1
Triglopsis quadricornis

109.58 14

20.44 14
Family Cyclopteridae
Cyclopteropsis jordani

109.64 14

23.23 14
Cyclopterus lumpus 12.7 9.5 95.03 1
Eumicrotremus spinosus

98.97 14

16.74 14
Family Hemitripteridae
Hemitripterus americanus

84.27 14

14.79 14
Family Liparidae
Liparis tunicatus

108.82 14

16.92 14
Family Platycephalidae
Platycephalus indicus

51.54 14

11.93 14
Family Scorpaenidae
Pterois volitans

45.79 14

10.47 14
Scorpaena cardinalis

87.20 14

25.53 14
Scorpaenopsis oxycepgus

70.87 14

11.49 14
Family Sebastidae
Helicolenus barathri

65.95 14

13.36 14
Helicolenus percoides

70.46 14

12.93 14
Sebastes alutus

59.73 14

12.82 14
Sebastes atlanticus

81.20 14

14.78 14
Sebastes marinus 12.4 8.2 79.86 3



298 3
" " 8.6 6.4 42.80 1
Sebastes polyspinis

58.58 14

17.36 14
Family Triglidae
Chelidonichthys cuculus 10.3 7.9 64.37 1
Chelidonichthys gurnardus 11.1 8.5 73.89 1
Chelidonichthys lucerna 9.5 7.1 53.46 1
Prionotus carolinus 9.2 6.5 46.97 4 4.3 3.2 10.81 4
" " 11.2 7.3 64.21 7 4.8 3.0 11.31 7
Prionotus evolans 10.0 57.0 447.68 4 4.7 2.7 9.97 4
ORDER Siluriformes
Family Callichthyidae
Corydoras metae

98.71 14

25.09 14
Family Ictaluridae
Ameiurus catus 10.4 8.7 71.06 3



123 3
Family Loricariidae
Farlowella acus

84.53 14

15.65 14
Liposarcus pardalis

113.83 14

11.43 14
Hypostomus plecostomus

79.06 14

19.84 14
Family Mochokidae
Synodontis notatus

55.95 14

10.27 14
" " (triploid?)

74.05 14

13.50 14
Family Pimelodidae
Pimelodella gracilis 15.0 9.0 106.03 9 4.5 3.0 10.60 9
Family Plotosidae
Euristhmus lepturus

114.00 14

21.90 14
ORDER Stomiiformes
Family Stomiidae
Idiacanthus atlanticus

218.07 14

21.71 14
ORDER Syngnathiformes
Family Centriscidae
Aeoliscus strigatus

52.75 14

7.74 14
Centriscops humerosus

81.62 14

14.36 14
Family Fistulariidae
Fistularia petimba

51.76 14

12.93 14
Family Syngnathidae
Corythoichthys intestinalis

101.04 14

25.42 14
Hippocampus abdominalis

81.26 14

14.44 14
Syngnathus fuscus 10.6 9.1 75.76 6
Syngnathus scovelli

46.96 14

5.46 14
Syngnathus typhle 11.1 9.5 83.14 1
ORDER Tetraodontiformes
Family Balistidae
Abalistes stellatus

43.91 14

7.36 14
Balistes carolinensis 9.2 6.6 47.69 13 4.0 2.7 8.48 13 127 13
Balistapus undulatus

54.48 14

9.14 14
Sufflamen fraenatus

43.92 14

11.49 14
Family Molidae
Mola mola 12.5 9.3 91.30 4 5.1 4.5 18.02 4
Family Monacanthidae
Aluterus schoepfii 8.2 7.2 46.37 13 3.9 2.9 8.88 13 181 13
Pseudomonacanthus peroni

35.43 14

10.53 14
Family Tetraodontidae
Arothron manilensis

46.74 14

9.47 14
Canthigaster bennetti

43.86 14

6.94 14
Canthigaster valentini

40.26 14

5.60 14
Lagocephalus lunaris

50.91 14

7.30 14
" " (triploid?)

58.24 14

12.98 14
Sphoeroides maculatus 8.8 6.9 47.69 4 3.4 2.8 7.48 4
Tetraodon nigroviridis

46.12 14

8.46 14
Family Triacanthidae
Tripodichthys angustifrons

36.51 14

8.03 14
ORDER Zeiformes
Family Oreosomatidae
Neocyttus rhomboidalis

213.83 14

27.68 14
Family Zeidae
Zenopsis nebulosus

100.99 14

23.88 14
Zeus faber 12.7 9.5 95.03 1
SUBCLASS SARCOPTERYGII
INFRACLASS Dipnoi
ORDER Ceratodontiformes
Family Ceratodontidae
Neoceratodus forsteri 39.0 24.0 735.13 2 14.0 9.8 107.76 2 5700 8
ORDER Lepidosireniformes
Family Protopteridae
Protopterus annectens annectens 44.6 27.0 944.70 1 17.5 8.8 120.09 1

References: 1) Gulliver 1875, 2) Cleland and Johnston 1912, 3) Wintrobe 1933, 4) Kisch 1949a, 5) Kisch 1949b, 6) Kisch 1951, 7) Hartman and Lessler 1964, 8) Pedersen 1971, 9) Conroy and Rodriguez 1966, 10) Saunders 1966, 11) Potter et al. 1982, 12) Banerjee et al. 1988, 13) Lewis 1996, 14) Hardie and Hebert 2003.

Click here for the reference list.

Last Modified By: T. Ryan Gregory
Last Modified On: Thursday, October 16, 2003